A List of Butterflies from Japan

By Hirotaka MATSUKA
2003.6.10 (revised)

How many butterfly species are there in Japan?

Two hundreds and thirty-seven species are listed as native ones here.
The categories used in this list are summarized as follows:

1) Native species: Butterflies regularly breeding in Japan. Immigrant species appearing continuously for more than 20 years, and also the naturalized species were included in the list from 2001. ---237 species.

2) Immigrant species: Butterflies invading from neighboring countries by natural causes like a typhoon and etc. ---63 species

3) Deliberate introduction species: Butterflies recorded by an artificial primary factor intentionally, or irrespective of unconsciousness--- 6 species

4) Status uncertain species--- 12 species

5) Doubtful species: The records of the butterflies apparently are errors or falsehood. They should be removed from list of Japanese butterflies list--- 32 species.

Contents

Abbreviations of locality

Hesperiidae 36 species
Papilionidae 21 species
Pieridae 23 species
Lycaenidae 70 species
Libytheidae 1 species
Danaidae 5 species
Nymphalidae 53 species
Satyridae 28 species

[Immigrant species] 63 species
[Deliberate introduction species] 6 species
[Status uncertain species] 12 species
[Doubtful species] 32 species

Abbreviations of locality [Map of Japan] [Contents]

Am ---Amami (Is.)
HK --- Hokkaido
HN --- Honshu
Ir ---- Iriomotejima (I.)
Is ---- Ishigakijima (I.)
KY --- Kyushu
Mi --- Miyako (Is.)
Og --- Ogasawara (Is.)
Ok --- Okinawa (Is.)
Oki--- Oki (Is.)
SI ---- Sikoku
Ta --- Tanegasima (I.)
To --- Tokara (Is.)
Ts --- Tsusima (I.)
Ya ----Yakushima (I.)
Yo ----Yonagunijima (I.)
< > ---- may not be native.
Red letters ---- The subspecies are described double.

Hesperiidae [Contents]

Pyrgus maculatus (Bremer & Grey, 1852) subsp. maculatus (Bremer & Grey, 1852) HK HN SI
Pyrgus malvae (Linnaeus, 1758) HK
Erynnis montanus (Bremer, 1861) HK HN SI KY
Daimio tethys (Menetries, 1857) subsp. tethys (Menetries, 1857) HK HN SI KY Ts
Tagiades trebellius (Hopffer, 1874) subsp. martinus Plotz, 1884 Is Ir
Bibasis aquilina (Speyer, 1879) subsp. chrysaeglia (Butler, [1882]) HK HN SI KY Ts
Choaspes benjaminii (Guerin-Meneville, 1843)
--------subsp. japonica (Murray, 1875) HN SI KY Ts Ta Ya Am Ok
--------subsp. formosana (Fruhstorfer, 1911) Is Ir
Badamia exclamationis (Fabricius, 1775) Is Ir
Hasora chromus (Cramer, [1780]) subsp. inermis Elwes & Edwards, 1897 Am Ok Is Ir Yo
Hasora badra (Moore, [1858]) subsp. badra (Moore, [1858]) Ir <Yo> [Immigrant species from 1974]
Erionota torus Evans, 1941 Ok [Deliberate introduction species from 1971]
Leptalina unicolor (Bremer & Grey, 1852) HK HN SI KY
Carterocephalus palaemon (Pallas, 1771)
--------subsp. satakei (Matsumura, 1919) HN
--------subsp. akaishianus Fujioka, 1970 HN
Carterocephalus sylvicola (Meigen, 1829) HK
Aeromachus inachus (Menetries, 1858) subsp. inachus (Menetries, 1858) HN Ts
Thoressa varia (Murray, 1875) HK HN SI KY
Isoteinon lamprospilus C. & R. Felder, 1862 subsp. lamprospilus C. & R. Felder, l862 HN SI KY Ts
Thymelicus leoninus (Butler, 1878)
--------subsp. leoninus (Butler, 1878) HK HN KY
--------subsp. hamadakohi Fujioka, 1993 SI
Thymelicus sylvaticus (Bremer, 1861) subsp. sylvaticus (Bremer, 1861) HK HN SI KY
Ochlodes ochraceus (Bremer, 1861) HK HN SI KY
Ochlodes venatus (Bremer & Grey, 1852) subsp. venatus (Bremer & Grey, 1852) HK HN
Ochlodes asahinai Shirozu, 1964 Is Ir
Hesperia florinda (Butler, 1878) HN
Potanthus flavus (Murray, 1875) subsp. flavus (Murray, 1875) HK HN SI KY Ts Ta Ya
Telicota colon (Fabricius, 1775) subsp. stinga Evans, 1949 Is Ir Yo
Notocrypta curvifascia (C. & R. Felder, 1862) subsp. curvifascia (C. & R. Felder, 1862) HN KY Ta Ya To Am Ok Is Ir Yo
Udaspes folus (Cramer, [1775]) Am Mi Is Ir Yo
Polytremis pellucida (Murray, 1875) subsp. pellucida (Murray, 1875) HK HN SI KY Ts
Pelopidas jansonis (Butler, 1878) HN SI KY
Pelopidas mathias (Fabricius, 1798) subsp. oberthueri Evans, 1937 HN SI KY Ts Ta Ya To Am Ok Mi Is Ir Yo
Pelopidas agna (Moore, [1866]) Ok Mi Is Ir Yo
Parnara ogasawarensis Matsumura, 1906 Og
Parnara guttata (Bremer & Grey, 1852) subsp. guttata (Bremer & Grey, 1852) HK HN SI KY Ta Ya To Am Ok Mi Is Ir Yo
Parnara naso (Fabricius, 1798) subsp. bada (Moore, 1878) Am Ok Is Ir Yo
Borbo cinnara (Wallace, 1866) Am Ok Mi Is Ir Yo
Suastus gremius (Fabricius, 1798) subsp. gremius (Fabricius, 1798) <Am> Ok Mi Is Ir <Yo> [Deliberate introduction species from 1973]

Papilionidae [Contents]

Parnassius glacialis Butler, 1866 HK HN SI
Parnassius stubbendorfii Menetries, 1849 subsp. hoenei Schweitzer, 1912 HK
Parnassius eversmanni Menetries, 1855 subsp. daisetsuzanus Matsumura, 1926 HK
Luehdorfia japonica Leech, 1889 HN
Luehdorfia puziloi (Erschoff, 1872)
--------subsp. yessoensis Rothschild, 1918 HK
--------subsp. inexpecta Sheljuzhko, 1913 HN
Byasa alcinous (Klug, 1836)
--------subsp. alcinous (Klug, 1836) HN SI KY Ts
--------subsp. yakushimana (Esaki & Umeno, 1929) Ta Ya
--------subsp. loochooana (Rothschild, 1896) Am Ok
--------subsp. miyakoensis Omoto, 1960 Mi
--------subsp. bradanus (Fruhstorfer, 1908) Is Ir Yo
Pachliopta aristolochiae (Fabricius, 1775) subsp. interpositus (Fruhstorfer, 1902) <Am Ok> Mi Is Ir [Immigrant species from 1968]
Sericinus montela (Bremer & Grey, 1852) HN [Deliberate introduction species from 1978]
Graphium sarpedon (Linnaeus, 1758) subsp. nipponum (Fruhstorfer, 1903) HN SI KY Ts Ta Ya Am Ok Mi Is Ir Yo
Graphium doson (C. & R. Felder, 1864)
-------- subsp. albidum (Wileman, 1903) HN SI KY Ts Ta Ya Am Ok
--------subsp. perillus (Fruhstorfer, 1908) Is Ir
Papilio xuthus Linnaeus, 1767 HK HN SI KY Ts Ta Ya Am Ok Mi Is Ir Yo
Papilio machaon Linnaeus, 1758 subsp. hippocrates C. & R. Felder, 1864 HK HN SI KY Ts Ta Ya
Papilio polytes Linnaeus, 1758 subsp. polytes Linnaeus, 1758 Am Ok Mi Is Ir Yo
Papilio helenus Linnaeus, 1758 subsp. nicconicolens Butler, 1881 HN SI KY Ts Ta Ya To Am Ok Mi Is Ir Yo
Papilio protenor Cramer, [1775]
--------subsp. demetrius Stoll, [1782] HN SI KY Am
--------subsp. liukiuensis Fruhstorfer, [1899] Ok Mi Is Ir Yo
Papilio macilentus Janson, 1877 HK HN SI KY
Papilio memnon Linnaeus, 1758
--------subsp. thunbergii von Siebold, 1824 HN SI KY Ta Ya To Am Ok Mi Is Ir Yo
--------subsp. heronus Fruhstorfer, 1902 <Immigrant subsp.>
Papilio dehaanii C. & R. Felder, 1864
--------subsp. dehaanii C. & R. Felder, 1864 HK HN SI KY
--------subsp. hachijonis Matsumura, 1919 HN
--------subsp. tokaraensis Fujioka, 1975 To
--------subsp. amamiensis (Fujioka, 1981) Am
--------subsp. takasago Nakahara & Esaki, 1930 <Immigrant subsp.>
Papilio okinawensis Fruhstorfer, 1898 Ok
Papilio junia Jordan, 1909 Is Ir Yo
Papilio maackii Menetries, 1858 HK HN SI KY Ts Ta Ya

Pieridae [Contents]

Leptidea amurensis (Menetries, 1858) HK HN KY
Leptidea morsei (Fenton, [1882]) HK
Eurema hecabe (Linnaeus, 1758) subsp. hecabe (Linnaeus, 1758) HN SI KY Og Ts Ta Ya To Am Ok Mi Is Ir Yo
Eurema blanda (Boisduval, 1836) subsp. arsakia (Fruhstorfer, 1910) Is Ir Yo
Eurema laeta (Boisduval, 1836) subsp. betheseba (Janson, 1878) HN SI KY
Colias erate (Esper, [1805]) subsp. poliographus Motschulsky, 1860 HK HN SI KY Og Ts Ta Ya To Am Ok Mi Is Ir Yo
Colias palaeno (Linnaeus, 1761)
--------subsp. aias Fruhstorfer, 1903 HN
--------subsp. sugitanii Esaki, 1929 HN
Gonepteryx aspasia (Menetries, 1858) subsp. niphonica Bollow, 1930 HN SI KY
Gonepteryx rhamni (Linnaeus, 1758) subsp. maxima Butler, 1885 HN
Catopsilia pyranthe (Linnaeus, 1758) subsp. pyranthe (Linnaeus, 1758) To Ok Mi Is Ir Yo
Catopsilia pomona (Fabricius, 1775) subsp. pomona (Fabricius, 1775) Og KY Ta Ya To Am Ok Mi Is Ir Yo
Aporia hippia (Bremer, 1861) subsp. japonica Matsumura, 1919 HN
Aporia crataegi (Linnaeus, 1757) subsp. adherbal Fruhstorfer, 1910 HK
Artogeia melete (Menetries, 1857)
--------subsp. melete (Menetries, 1857) HK HN SI KY Ts Ta Ya Am
Artogeia dulcinea (Butler, 1882)
--------subsp. pseudonapi (Verity , �m1911�n) HK (eastern)
Artogeia nesis (Fruhstorfer, 1909) HK (Southern) HN SI KY
Artogeia rapae (Linnaeus, 1758)
--------subsp. crucivora (Boisduval, 1836) HK HN SI KY Og Ts Ta Ya To Am Ok Mi Is Ir Yo
Artogeia canidia (Sparrman, 1768)
--------subsp. juba (Fruhstorfer, 1910) Ts
--------subsp. canidia (Sparrman, 1768)  <Immigrant subsp.>
Anthocharis scolymus Butler, 1866 HK HN SI KY Ta Ya
Anthocharis cardamines (Linnaeus, 1758)
--------subsp. isshikii Matsumura, 1925 HN
--------subsp. hayashii Fujioka, 1970 HN
Appias paulina (Cramer, [1777]) subsp. minato (Fruhstorfer, [1899]) To Am Ok Mi Is Ir Yo
Appias lyncida (Cramer, [1777]) subsp. formosana (Wallace, 1866) <Ir> Yo [Immigrant species from 1966]
Hebomoia glaucippe (Linnaeus, 1758) subsp. liukiuensis Fruhstorfer, 1898 KY Ta Ya To Am Ok Mi Is Ir Yo

Lycaenidae [Contents]

Panchala ganesa (Moore, [1858]) subsp. loomisi (H. Pryer, 1886) HN SI KY Ya
Narathura japonica (Murray, 1875) subsp. japonica (Murray, 1875) HN SI KY Ts Ta Ya To Am Ok Mi Is Ir
Narathura bazalus (Hewitson, 1862) subsp. turbata (Butler, [1882]) HN SI KY Og Ts Ta Ya To Am Ok Is Ir Yo
Artopoetes pryeri (Murray, 1873) HK HN SI KY
Ussuriana stygiana (Butler, 1881) HK HN SI KY
Coreana raphaelis (Oberthur, 1880) subsp. yamamotoi Okano, 1953 HN
Shirozua jonasi (Janson, 1877) HK HN
Japonica lutea (Hewitson, [1865]) subsp. lutea (Hewitson, [1865]) HK HN SI KY Ts
Japonica onoi Murayama, 1953
--------subsp. onoi Murayama, 1953 HK HN
--------subsp. mizobei Saigusa, 1993 HN
Japonica saepestriata (Hewitson, [1865])
--------subsp. saepestriata (Hewitson, [1865]) HK HN SI
--------subsp. gotohi Saigusa, 1993 HN
Iratsume orsedice (Butler, [1882]) subsp. orsedice (Butler, [1882]) HK HN SI KY
Wagimo signatus (Butler, [1882]) HK HN KY SI
Araragi enthea (Janson, 1877) subsp. enthea (Janson, 1877) HK HN SI KY
Antigius attilia (Bremer, 1861) subsp. attilia (Bremer, 1861) HK HN SI KY Ts
Antigius butleri (Fenton, [1882]) HK HN KY
Neozephyrus japonicus (Murray, 1875) HK HN SI KY
Chrysozephyrus brillantinus (Staudinger, 1887) HK HN SI KY
Chrysozephyrus smaragdinus (Bremer, 1861) subsp. smaragdinus (Bremer, 1861) HK HN SI KY
Chrysozephyrus hisamatsusanus (Nagami & Ishiga, 1935) HN SI KY
Chrysozephyrus ataxus (Westwood, [185l])
--------subsp. kirishimaensis (Okajima, 1922) HN SI KY Ts
--------subsp. yakushimaensis (Yazaki, [1924]) Ya
Favonius yuasai Shirozu, 1947 HN KY
Favonius orientalis (Murray, 1875) HK HN SI KY
Favonius jezoensis (Matsumura, 1915) HK HN SI KY
Favonius taxila (Bremer, 1861) HK HN
Favonius ultramarinus (Fixsen, 1887) HK HN KY
Favonius cognatus (Staudinger, 1892) subsp. latifasciatus Shirozu & Hayashi, 1959 HN
Favonius saphirinus (Staudinger, 1887) HK HN SI KY Ts
Sibataniozephyrus fujisanus (Matsumura, 1910) HK HN SI KY
Strymonidia w-album (Knoch,1782) subsp. fentoni (Butler, [1882]) HK HN SI KY
Strymonidia iyonis Ota & Kusunoki, 1957
--------subsp. surugaensis Fujioka, 1981 HN
--------subsp. kiiensis Hara, 1984 HN
--------subsp. kibiensis Shirozu & M. Nanba, 1973 HN
--------subsp. iyonis Ota & Kusunoki, 1957 SI
Strymonidia mera (Janson, 1877) HK HN SI KY
Strymonidia pruni (Linnaeus, 1758) subsp. jezoensis (Matsumura, 1919) HK
Callophrys ferrea (Butler, 1866) HK HN SI KY
Rapala arata (Bremer, 1861) HK HN SI KY Ts
Deudorix eryx (Linnaeus, 1771) subsp. okinawana (Matsumura, 1919) Am Ok Is Ir
Spindasis takanonis (Matsumura, 1906) HN
Taraka hamada (H.Druce, 1875) subsp. hamada (H. Druce, 1875) HK HN SI KY Ts Ta Ya
Lycaena phlaeas (Linnaeus, 1761) subsp. daimio (Matsumura, 1919) HK HN SI KY Ts Ta Ya
Niphanda fusca (Bremer & Grey, 1852) HN SI KY Ts
Euchrysops cnejus (Fabricius, 1798) Am Ok Mi Is Ir Yo
Lampides boeticus (Linnaeus, 1767) HN SI KY Am Ok Mi Is Ir Yo
Nacaduba kurava (Moore, [1858]) subsp. septentrionalis Shirozu, 1953 KY Am Ok Mi Is Ir Yo
Prosotas nora (C.Felder, 1860) subsp. kanoi (Omoto, 1959) Is Ir Yo
Jamides bochus (Stoll, [1782]) subsp. ishigakianus Shirozu, 1953 <KY>Is Ir Yo
Zizeeria maha (Kollar, [1844])
--------subsp. argia (Menetries, 1857) HN SI KY To
--------subsp. okinawana (Matsumura, 1929) To Am Ok Mi Is Ir Yo
Zizeeria karsandra (Moore, 1865) Ok Mi Is Ir Yo
Zizina otis (Fabricius, 1787)
--------subsp. emelina (de l'Orza, 1869) HN SI KY Ts Ta
--------subsp. riukuensis (Matsumura, 1929) To Am Ok Mi Is Ir Yo
Glaucopsyche lycormas (Butler, 1866) subsp. lycormas (Butler, 1866) HK HN
Maculinea teleius (Bergstrasser, 1779)
--------subsp. ogumae (Matsumura, 1910) HK
--------subsp. matsumurai Shirozu, 1953 HK
--------subsp. shiriyensis (Matsumura, 1927) HN
--------subsp. kazamoto (H. Druce, 1875) HN KY
--------subsp. hosonoi A. Takahashi, 1973 HN
--------subsp. hakusanensis Fujioka, 1975 HN
Maculinea arionides (Staudinger, 1887) subsp. takamukui (Matsumura, 1919) HK HN
Shijimiaeoides divinus (Fixsen, 1887) subsp. barine (Leech, [1893]) HN KY
Megisba malaya (Horsfield, [1828]) subsp. sikkima Moore, 1884 Am Ok Mi Is Ir Yo
Acytolepis puspa (Horsfield, [1828]) subsp. ishigakiana (Matsumura, 1929) HN SI KY Ta Ya Am Ok Is Ir
Udara albocaerulea (Moore, 1879) subsp. albocaelulea (Moore, 1879) HN SI KY Ts Ta Ya To Am Ok
Celastrina argiolus (Linnaeus, 1758) subsp. ladonides (de l'Orza, 1869) HK HN SI KY Ts Ta Ya To Am
Celastrina sugitanii (Matsumura, 1919)
--------subsp. ainonica Murayama, 1952 HK
--------subsp. sugitanii (Matsumura, 1919) HN SI
--------subsp. kyushuensis Shirozu, 1943 KY
Celastrina ogasawaraensis (H.Pryer, 1883) Og
Vacciniina optilete (Knoch, 1781) subsp. daisetsuzana (Matsumura, 1926) HK
Scolitantides orion (Pallas, 1771) subsp. jezoensis (Matsumura, 1919) HK
Tongeia fischeri (Eversmann, 1843) HN SI KY Ts
Everes argiades (Pallas, 1771) subsp. hellotia (Menetries, 1857) HK HN SI KY Ts Ta Ya Am To <Yo>
Everes lacturnus (Godart, [1824])
--------subsp. kawaii Matsumura, 1926 HN SI KY Ta Ya
--------subsp. rileyi Godfrey, 1916 To Am Ok Ir
Shijimia moorei (Leech, 1889) subsp. moorei (Leech, 1889) HN KY
Pithecops corvus Fruhstorfer, 1919 subsp. ryukyuensis Shirozu, 1964 Ok Is Ir
Pithecops fulgens Doherty, 1889 subsp. tsushimanus Shirozu & Urata, 1957 Ts
Neopithecops zalmora (Butler, 1870) Ir Is
Plebejus argus (Linnaeus, 1758)
--------subsp. pseudaegon (Butler, [1882]) HK
--------subsp. micrargus (Butler, 1878) HN KY
Lycaeides argyrognomon (Bergstrasser, 1779) subsp. praeterinsularis (Verity, 1921) HN
Lycaeides subsolanus (Eversmann, 1851)
--------subsp. iburiensis (Butler, [1882]) HK
--------subsp. yaginus (Strand, 1922) HN
--------subsp. yarigadakeanus (Matsumura, 1929) HN
Curetis acuta Moore, 1877 subsp. paracuta de Niceville, 1901 HN SI KY Ts Ta Ya To Am Is Ir

Libytheidae [Contents]

Libythea celtis (Laicharting, 1782)
--------subsp. celtoides Fruhstorfer, [1909] <HK> HN SI KY Og Ta Ya
--------subsp. amamiana Shirozu, 1956 Am Ok Is
--------subsp. formosana Fruhstorfer, 1908 <Immigrant subsp.>

Danaidae [Contents]

Anosia chrysippus (Linnaeus, 1758) subsp. chrysippus (Linnaeus, 1758) Am Ok Mi Is Ir Yo
Salatura genutia (Cramer, [1779]) subsp. genutia (Cramer, [1779]) Mi Is Ir Yo
ldea leuconoe Erichson, 1834
--------subsp. clara (Butler, 1867) Am Ok
--------subsp. riukiuensis (Holland, 1893) Mi Is Ir Yo
Radena similis (Linnaeus, 1758) subsp. similis (Linnaeus, 1758) Am Ok Mi Is Ir Yo
Parantica sita (Kollar, [1844]) subsp. niphonica (Moore, 1883) HN SI KY Ts Ta Ya To Am Ok Mi Is Ir Yo

Nymphalidae [Contents]

Mellicta ambigua Menetries, 1859 subsp. niphona (Butler, 1878) HN
Melitaea regama Fruhstorfer, 1915 HN
Melitaea scotosia Butler, 1878 HN
Clossiana freija (Thunberg, 1791) subsp. asahidakeana (Matsumura, 1926) HK
Clossiana iphigenia (Graeser, 1888) HK
Clossiana thore (Hubner, [1803-1804]) subsp. jezoensis (Matsumura, 1919) HK
Brenthis daphne ([Denis & Schiffermuller], 1775)
--------subsp. iwatensis (Okano, 1951) HK HN
--------subsp. rabdia (Butler, 1877) HN
Brenthis ino (Rottemburg, 1775)
--------subsp. mashuensis (Kono, 1931�j HK
--------subsp. tigroides (Fruhstorfer, 1907) HN
--------subsp. tanigawensis (Nakahara, 1941) HN
Argyronome laodice (Pallas, 1771) subsp. japonica (Menetries, 1857) HK HN SI KY Ts
Argyronome ruslana (Motschulsky, 1866) subsp. lysippe (Janson, 1877) HK HN SI KY Ts
Damora sagana (Doubleday, [1847]) subsp. ilone (Fruhstorfer, 1907) HK HN SI KY Ts <Yo>
Nephargynnis anadyomene (C. & R. Felder, 1862) subsp. midas (Butler, 1866) HK HN SI KY Ts
Argynnis paphia (Linnaeus, 1758) subsp. tsushimana Fruhstorfer, 1906 HK HN SI KY Ts
Speyeria aglaja (Linnaeus, 1758)
--------subsp. basalis (Matsumura, 1908) HK
--------subsp. fortuna (Janson, 1877) HN
Fabriciana adippe ([Denis & Schiffermuller], 1775)
--------subsp. pallescens (Butler, 1873) HK HN SI KY Ts
Fabriciana niobe Linnaeus, 1758 subsp. tsubouchii Fujioka, 2002 HK
Fabriciana nerippe (C. & R. Felder, 1862) HN SI KY
Argyreus hyperbius (Linnaeus, 1763) HN SI KY Ta Ya To Am Ok Mi Is Ir Yo
Cupha erymanthis (Drury, 1773) <Is> Ir [Immigrant species from 1973]
Athyma selenophora (Kollar, [1844]) subsp. ishiana Fruhstorfer, 1899 Is Ir
Athyma perius (Linnaeus, 1758) subsp. perius (Linnaeus, 1758) <Is> Ir Yo [Immigrant species from 1967]
Ladoga camilla (Linnaeus, 1764) subsp. japonica (Menetries, 1857) HK HN SI KY
Ladoga glorifica (Fruhstorfer, 1909) HN
Limenitis populi (Linnaeus, 1758) subsp. jezoensis Matsumura, 1919 HK HN
Neptis sappho (Pallas, 1771) subsp. intermedia W. B. Pryer, 1877 HK HN SI KY Ts Ta Ya
Neptis hylas (Linnaeus, 1758) subsp. luculenta Fruhstorfer, 1907 Am Ok Mi Is Ir Yo
Neptis pryeri Butler, 1871
--------subsp. Fukuda, Minotani et Iwano, 2000 HN
--------subsp. iwasei Fujioka, 1999 HN
--------subsp. setoensis Fukuda et Minotani, 1999 HN SI KY
--------subsp. hamadai Fujioka, Minotani et Fukuda, 1999 SI
--------subsp. yodoei Fujioka, 1999 Oki
Neptis rivularis (Scopoli, 1763)
--------subsp. bergmanni Bryk, 1942 HK
--------subsp. shirozui Okano, 1954 HN
--------subsp. tadamiensis Higuma, 1961 HN
--------subsp. insularum Fruhstorfer, 1907 HN
Neptis philyra Menetries, 1858 subsp. excellens Butler, 1878 HK HN SI KY Ts
Neptis alwina (Bremer & Grey, 1852) HK HN
Araschnia burejana Bremer, 1861 subsp. strigosa Butler,1866 HK HN SI KY
Araschnia levana (Linnaeus, 1758) subsp. obscura Fenton, [1882] HK
Polygonia c-aureum (Linnaeus, 1758) HK HN SI KY
Polygonia c-album (Linnaeus, 1758) subsp. hamigera (Butler, 1877) HK HN SI KY
Nymphalis xanthomelas ([Denis & Schiffermuller], 1775) subsp. japonica (Stichel, 1902) HK HN SI KY
Nymphalis vaualbum ([Denis & Schiffermuller], 1775) subsp. samurai (Fruhstorfer, 1907) HK HN
Nymphalis antiopa (Linnaeus, 1758) subsp. asopos (Fruhstorfer, 1909) HK HN
Inachis io (Linnaeus, 1758) subsp. geisha (Stichel, 1908) HK HN
Aglais urticae (Linnaeus, 1758) subsp. connexa (Butler, 1882) HK HN
Kaniska canace (Linnaeus, 1763)
--------subsp. nojaponicum (von Siebold, 1824) HK HN SI KY Ts Ta Ya
--------subsp. ishima (Fruhstorfer, [1899]) To Am Ok Mi Is Ir Yo
Vanessa indica (Herbst, 1794) HK HN SI KY Og Ts Ta Ya To Am Ok Mi Is Ir Yo
Cynthia cardui (Linnaeus, 1758) HN SI KY Am Ok Mi Is Ir Yo
Junonia almana (Linnaeus, 1758) subsp. almana (Linnaeus, 1758) KY Ta Ya To Am Ok Mi Is Ir Yo
Junonia orithya (Linnaeus, 1758) subsp. orithya (Linnaeus, 1758) Is Ir Yo
Kallima inachus (Doyere, [1840]) subsp. eucerca Fruhstorfer, 1898 Am Ok Is Ir
Hypolimnas misippus (Linnaeus, 1764) Is Ir
Dichorragia nesimachus (Doyere, [1840])
--------subsp. nesiotes Fruhstorfer, 1903 HN SI KY Ta Ya
--------subsp. okinawaensis Shimagami, 1986 Am Ok
--------subsp. ishigakianus Shirozu, 1952 Is Ir
Cyrestis thyodamas Doyere, [1840] subsp. mabella Fruhstorfer, 1898 HN SI KY Ta Ya To Am Ok Is Ir Yo
Polyura eudamippus (Doubleday, 1843) subsp. weismanni (Fritze, 1894) Ok
Apatura metis Freyer, [1829] subsp. substituta Butler, 1873 HK HN SI KY Ts
Hestina japonica (C. & R. Felder, 1862) subsp. japonica (C. & R. Felder, 1862) HK HN SI KY Ts
Hestina assimilis (Linnaeus, 1758) subsp. shirakii Shirozu, 1955 Am
--------subsp. assimilis (Linnaeus, 1758) <Deliberate introduction species>
Sasakia charonda (Hewitson, [1863] subsp. charonda (Hewitson, [1863]) HK HN SI KY

Satyridae [Contents]

Ypthima argus Butler, 1866 HK HN SI KY Ya
Ypthima multistriata Butler, 1883
--------subsp. niphonica Murayama, 1969 HN Ta SI KY Ta Ya
--------subsp. tsushimana Murayama, 1969 Ts
Ypthima riukiuana Matsumura, 1906 Ok
Ypthima masakii Ito, 1947 Is Ir
Ypthima yayeyamana Nire, 1920 Is Ir
Erebia niphonica Janson, 1877
--------subsp. scoparia Butler, [1882] HK
--------subsp. niphonica Janson, 1877 HN
Erebia ligea (Linnaeus, 1758)
--------subsp. rishirizana Matsumura, 1928 HK
--------subsp. takanonis Matsumura, 1909 HN
Oeneis norna (Thunberg, 1791)
--------subsp. asamana Matsumura, 1919 HN
--------subsp. sugitanii Shirozu, 1952 HN
Oeneis melissa (Fabricius, 1775) subsp. daisetsuzana Matsumura, 1926 HK
Coenonympha oedippus (Fabricius, 1787)
--------subsp. annulifer Butler, 1877 HN
--------subsp. arothius Okada & Torii, 1945 HN
Coenonympha hero (Linnaeus, 1761)
--------subsp. latifasciata Matsumura, 1925 HK
--------subsp. neoperseis Fruhstorfer, 1908 HK
Minois dryas (Scopoli, 1763) subsp. bipunctata (Motschulsky, 1860) HK HN SI KY Ts
Lopinga achine (Scopoli, 1763)
--------subsp. jezoensis (Matsumura, 1919) HK
--------subsp. oniwakiensis Y. Yazaki & Hiramoto, 1981 HK
--------subsp. achinoides (Butler, 1878) HN
Lasiommata deidamia (Eversmann, 1851)
--------subsp. deidamia (Eversmann, 1851) HK
--------subsp. interrupta (Fruhstorfer, 1909) HN
--------subsp. kampuzana Y. Yamazaki, 1981 SI
Zophoessa callipteris (Butler, 1877) HK HN SI KY
Kirinia epaminondas (Staudinger, 1887) HK HN SI KY
Ninguta schrenckii (Menetries, 1858) HK HN
Lethe sicelis (Hewitson, [1862]) HN SI KY
Lethe diana (Butler, 1866)
--------subsp. diana (Butler, 1866) HK HN SI KY Ts
--------subsp. mikuraensis Shirozu 1975 HN
Lethe marginalis (Motschulsky, 1860) HN SI KY
Lethe europa (Fabricius, 1775) subsp. pavida Fruhstorfer, 1908 Is Ir Yo
Neope goschkevitschii (Menetries, 1857) HK HN SI KY
Neope niphonica Butler, 1881
--------subsp. niphonica Butler, 1881 HK HN SI KY
--------subsp. kiyosumiensis M. Takahashi & Aoyama, 1981 HN
--------subsp. marumoi Esaki & Umeno, 1929 Ya
Mycalesis gotama Moore, 1858 subsp. fulginia Fruhstorfer, 1911 HK HN SI KY Ts Ta Ya
Mycalesis madjicosa Butler, 1868
--------subsp. amamiana Fujioka, 1975 Am Ok
--------subsp. madjicosa Butler, 1868 Is Ir Yo
Mycalesis francisca (Stoll, [1780]) subsp. perdiccas Hewitson, [1862] HN SI KY
Melanitis phedima (Cramer, 1780) subsp. oitensis Matsumura, 1919 HN SI KY Ts Ta Ya Am Ok
Melanitis leda (Linnaeus, 1758) subsp. ismene (Cramer, [1775]) Og Am Ok Mi Is Ir Yo

[Immigrant species] [Contents]

Hasora taminatus (Huebner, [1818]) subsp. vairacana Fruhstorfer, 1911
Graphium agamemnon (Linnaeus, 1758)
Papilio demoleus Linnaeus, 1758 subsp. libanius Fruhstorfer, 1908
Papilio alphenor Cramer, [1776] subsp. ledebouria Eschsholtz, 1821
Papilio castor Westwood, 1842 subsp. formosanus Rothschild, 1896
Papilio memnon Linnaeus, 1758
--------subsp. heronus Fruhstorfer, 1902
Papilio rumanzovia Eschscholtz, 1821
Papilio dehaanii C. & R. Felder, 1864
--------subsp. takasago Nakahara & Esaki, 1930
Eurema brigitta (Stoll, [1780]) subsp. hainana (Moore, 1878)
Catopsilia scylla (Linnaeus, 1763) subsp. cornelia Fabricius, 1787
Leptosia nina (Fabricius, 1793) subsp. niobe (Wallace, 1866)
Artogeia canidia (Sparrman, 1768)
--------subsp. canidia (Sparrman, 1768)
Pieris brassicae (Linnaeus, 1758) subsp. brassicae (Linnaeus, 1758)
Pontia daplidice (Linnaeus, 1758)
Appias albina (Boisduval, 1836) subsp. semperi (Moore, [1905])
Appias nero (Fabricius, 1793) subsp. domitia (C. & R. Felder, 1862)
Appias indra Moore, 1857
Saletara panda (Godart, 1819) subsp. nathalia (C. & R. Felder, 1862)
Ixias pyrene (Linnaeus, 1764) subsp. insignis Butler, 1879
Leptotes plinius (Fabricius, 1793)
Petrelaea tombugensis (Rober, 1886)
Jamides alecto (C. Felder, 1860) subsp. dromicus (Fruhstorfer, 1910)
Catochrysops panormus (C. Felder, 1860) subsp. exiguus (Distant, 1886)
Catochrysops strabo (Fabricius, 1793)
--------subsp. riama Corbet, 1938
--------subsp. luzonensis Tite, 1959
Zizula hylex (Fabricius, 1775)
Famegana alsulus (Herrich-Schaffer, 1869)
Udara dilecta (Moore, 1879) subsp. dilecta (Moore, 1879)
Celastrina lavendularis (Moore, 1877) subsp. himilcon (Fruhstorfer, 1909)
Chilades mindorus (C. & R. Felder, [1865])
Chilades pandava (Horsfield, [1829]) subsp. pandava (Horsfield, 1829)
Deudorix epijarbas (Moore, [1858])
Libythea celtis (Laicharting, 1782)
--------subsp. formosana Fruhstorfer, 1908
Libythea geoffroy Godart, [1824] subsp. philippina Staudinger, 1889
Salatura melanippus (Cramer, [1777]) subsp. edmondii (Lesson, 1837)
Tirumala septentrionis (Butler, 1874) subsp. septentrionis (Butler, 1874)
Tirumala limniace (Cramer, [1775])
--------subsp. limniace (Cramer, [1775])
--------subsp. orestilla (Fruhstorfer, [1910])
Tirumala hamata (MacLeay, [1826]) subsp. orientalis (Semper, 1879)
Tirumala ishmoides (Moore, 1883) subsp. sonitnus (Fruhstorfer, 1911)
Parantica swinhoei (Moore, 1883)
Parantica luzonensis (C. & R. Felder, 1863) subsp. luzonensis (C. & R. Felder, 1863)
Parantica aglea (Stoll, [1782]) subsp. maghaba (Fruhstorfer, 1909)
Ideopsis juventa (Cramer, 1777)
Euploea mulciber (Cramer, 1777)
--------subsp. barsine Fruhstorfer, 1904
--------subsp. dufresne Godat, 1823
Euploea camaralzeman Butler, 1866 subsp. cratis Butler, 1866
Euploea swainson (Godart, [1824]) subsp. swainson (Godart, [1824])
Euploea core (Cramer, [1780]) subsp. godartii Lucas, 1853
Euploea sylvester (Fabricius, 1793)
--------subsp. swinhoei Wallace, 1866
--------subsp. laetifica Butler, 1866
--------subsp. harrisii C. & R. Felder, 1865
Euploea midamus (Linnaeus, 1758)
Euploea tulliolus (Fabricius, 1793)
--------subsp. koxinga Fruhstorfer, 1908
--------subsp. polita Erichson, 1834
Euploea eunice (Quoy, 1815)
--------subsp. hobsoni (Butler, [1878])
--------subsp. eunice (Quoy, 1815)
Euploea klugii Moore, [1858] subsp. erichsonii C. & R. Felder, [1865]
Euploea diocletianus (Fabricius, 1793) subsp. diocletianus (Fabricius, 1793)
Ariadne ariadne (Linnaeus, 1763) subsp. pallidior (Fruhstorfer, 1899)
Phalanta phalantha (Drury, [1773])
Phalanta alcippe (Stoll, 1782)
Cirrochroa tyche C. & R. Felder, 1861
Parthenos sylla (Donovan, [1799]) subsp. philippensis Fruhstorfer, 1898
Junonia lemonias (Linnaeus, 1758) subsp. lemonias (Linnaeus, 1758)
Junonia hedonia (Linnaeus, 1764) subsp. ida (Cramer, 1775)
Junonia atlites (Linnaeus, 1763)
Junonia lintingensis (Osbeck, 1765)
Doleschallia polibete (Cramer, [1779]) subsp. philippensis Fruhstorfer, I912
Yoma sabina (Cramer, [1780]) subsp. podium Tsukada, 1985
Hypolimnas bolina (Linnaeus, 1758)
--------subsp. f.kezia (Butler, [1878])
--------subsp. f.philippensis (Butler, 1874)
--------subsp. f.jacintha (Drury, 1773)
--------subsp. f.bolina (Linnaeus, 1758)
Hypolimnas anomala (Wallace, 1869) subsp. truentus Fruhstorfer, [1913]
Timelaea albescens (Oberthur, 1886) subsp. formosana Fruhstorfer, 1908
Melanitis boisduvalia (C. & R. Felder, 1863)

[Deliberate introduction species] [Contents]

Caltoris cahira (Moore, 1877)
Thymelicus lineola (Ochsenheimer, 1808) HK
Calinaga budda Oberthur, 1881
Troides helena (Linnaeus, 1758)
Polyura narcaea (Hewitson, 1854)
Elymnias hypermnestra (Linnaeus, 1763) subsp. hainana Moore, 1878

[Status uncertain species] [Contents]

Troides aeacus Felder, 1860 subsp. kaguya (Nakahara et Esaki, 1930)
Colias fieldii Menetries, 1855
Gonepteryx amintha Blanchard, 1871 subsp. formosana (Furuhstorfer, 1908)
Talbotia naganum (Moore, 1884) subsp. karumii (Ikeda, 1937)
Narathura rama (Kollar, 1844)
Spalgis epius (Westwood, [1851]) subsp. dilama (Moore, 1878)
Nothodanis schaeffera (Eschscholtz, 1821) subsp. schaeffera (Eschscholtz, 1821)
Danaus plexippus (Linnaeus, 1758) subsp. plexippus (Linnaeus, 1758)
Parantica vitrina (C.& R. Felder, 1861)
Athyma fortuna (Leech,1889) subsp. kodahirai (Sonan, 1938)
Melanargia epimede (Staudinger, 1892)
Mycalesis perseus (Fabricius, 1775)

[Doubtful species] [Contents]

Parnassius bremeri Bremer, 1864
Parnassius nomion Fischer von Waldheim, 1823
Chilasa agestor (Gray, 1831)
Graphium cloanthus (Westwood, 1841)
Papilio nephelus Boisduval, 1836
Papilio paris Linnaeus, 1758
Delias hyparete (Linnaeus, 1758)
Appias libythea (Fabricius, 1775)
Pieris mannii (Mayer, 1851)
Strymonidia prunoides (Staudinger, 1887)
Phengaris atroguttata (Oberthur, 1876)
Curetis brunnea Wileman, 1909
Abisara schedeli Fruhstorfer, 1904
Clossiana selene ([Denis & Schiffermuller], 1775)
Araschnia prorsoides (Blanchard, 1871)
Limenitis helmanni Lederer, 1853
Aglais milberti (Godart, 1819)
Neptis soma Moore, 1858
Symbrenthia hippoclus (Cramer, l779)
Sephisa chandra Moore, 1857
Junonia iphita Cramer, 1779
Helcyra superba Leech, 1890
Ypthima baldus (Fabricius, I775)
Mycalesis sangaica Butler, 1877
Mycalesis mineus (Linnaeus, 1767)
Coenonympha arcania (Linnaeus, 1761)
Ochlodes subhyalina (Bremer & Grey, 1852)
Potanthus juno (Evans, 1932)
Celaenorrhinus asmara (Butler, 1879)
Pelopidas conjuncta (Herrich-Schaffer, 1869)
Pelopidas sinensis (Mabille, 1877)
Telicota augias (Linnaeus, 1767)

Hirotaka MATSUKA
Email: matsuka@tkd.att.ne.jp

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